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JACKSONVILLE, Ala. — Zerrick Cooper threw three touchdown passes to Shaq Davidson and Jacksonville State held on to beat East Tennessee State 34-27 on Saturday night in the first round of the FCS playoffs.

Jacksonville State (9-3) will play at seventh-seeded Maine (8-3) in the second round at noon Saturday on Morse Field at Alfond Stadium in Orono.

Cooper was 20 of 32 for 226 yards passing and is the first quarterback in the program to surpass 3,000 yards in a season. Davidson had five catches for 69 yards.

Austin Herink was 17 of 39 passing for 260 yards with a touchdown and interception for East Tennessee State, and his 10-yard run capped the scoring. It was the first playoff appearance for ETSU since 1996.

Trailing 34-27 late in the fourth quarter, Quay Holmes rushed for 20 yards and caught a 25-yard pass to help drive the Buccaneers to the Jacksonville State 4. But Zack Woodard forced Holmes to fumble the ball into the end zone for a touchback with 2:38 to play. ETSU got the ball back with 1:39 left and no timeouts, and drove to their own 44-yard line but the drive ended with an incomplete pass on fourth-and-10.

ETSU built a 20-10 halftime lead on two JJ Jerman field goals, a 3-yard TD toss from Herink to Evan Wick, and Titus Tuckers’s 60-yard interception return for a touchdown.

But it was all Jacksonville State for much of the second half.

Davidson made a juggling catch in the corner of the end zone from five yards out, and Cade Stinnett’s 22-yard field goal tied the game at 20 in the third quarter. In the fourth, Cooper threw a strike to Davidson in the middle of the field, who broke one tackle and stretched over the goal line for a 13-yard score. Marlon Bridges’ 32-yard pick-6 gave the Gamecocks a 34-20 lead with 10:35 remaining.

ETSU receiver Kobe Kelley was taken off the field on a stretcher late in the game. He gave a thumbs-up in response to the ovation from the crowd.